In a coded relationship, if M - N means M is the sister of N, M + N means M is the brother of N, and M × N means M is the daughter of N, what is the relationship of O if M - N + O?
Given that '?' represents greater than, '?' represents less than, '?' represents addition, and '†' represents subtraction, which of the following is true? 5 ? 3 ? 2 ? 4 † 1?